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Does Source drive need to be startup drive?

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I have 'cloned' my Imac g5 hard drive onto a Western Digital HD using Retrospect Duplicate. I have since made changes to my Imac HD and want to use Restrspect to update my external HD. Does the Source drive need to be the Startup drive or can I boot up on my external HD and and Duplicate my Imac HD onto my external HD? I am thinking it doesn't matter which drive you boot up on but not sure.

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